In the following passage some words have been deleted. Fill in the blanks with the help ofthe alternatives given.
Passage:
Science has made an (1)............... contribution to the relief of humansuffering, and humanity (2)............ a deep (3).................. of gratitude to scientists whose (4)................ and sacrifices have led to many (5)................ discoveries and inventions.
Fill in the blank (3) with the most appropriate option.
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